All species of Chara reproduce sexually and show highly advanced oogamy. The sex organs are the most distinctive features of the order Charales and are the most complicated among the thallophytes. Male and female gametangia are called antheridia and oogonia respectively. Male fructification (cluster of antheridia) is called globule and the female is nucule. They are borne at the nodes of short branches, globule towards lower side and nucule (female structure) towards upper side.

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